Dendroculus in Genshin Impact is part of patch 3.0 that brought the region of Sumeru to the Teyvat.
Collecting Dendroculus is part of the experience of exploring the new area and is quite a valuable one. You can stumble upon one of these items while checking a mountain, a cavern, or even a giant robot, and use them to earn great items that makes hunting Dendroculus an exciting idea.
Since there are a lot of Dendroculus in Genshin Impact, we have prepared this guide covering all the Dendroculus locations, including the ones added in Genshin Impact 3.6, with tips on how to get the more difficult ones.
On this page:
Explainers:
Dendroculus locations:
Version 3.0 ‘The Morn a Thousand Roses Brings’ Trailer | Genshin Impact
What Dendroculus are used for in Genshin Impact?
The Dendroculus are Sumeru’s Oculi – items you can find all over the region and deliver to a Statue of the Seven. While some Dendroculus are easy to spot, others are hidden or locked and only get accessible after completing quests, puzzles or acquiring specific skills.
There are a total of 110 Dendroculus in the region as of patch 3.0. Using them to level up Sumeru’s Statue of the Seven is a great way to earn important items, such as Primogems, Sumeru Shrine of Depths Keys, and flasks of Memory of Flourishing Green.
Version 4.0 is here! Our Fontaine guide can help with Fontaine’s Reputation and the Fountain of Lucine, and you can also redeem new codes for Primogems if you want Lyney – and check him out on our tier list. For improving Lyney, you’ll need lots of Rainbow Rose, and Lynette requires Lumidouce Bell.
You’ll also need luck from the Pity and 50/50 systems to Wish with Fate for them and future Banners, and you always need the right Adventure Rank for events.
Elsewhere, we have tips and tricks for beginners, Dendroculus locations, fishing locations, and TCG card locations.
Gandharva Ville and East of Devantaka Mountain Dendroculus locations in Genshin Impact
Here are all the Dendroculus locations in Gandharva Ville and East of Devantaka Mountain in Genshin Impact:
Dendroculus 1
Dendroculus 2
At the top of the tree in the middle of the tree’s leaves.
Dendroculus 3
Completing the Agnihotra Sutra World Quest is needed to open the path.
Dendroculus 4
It’s inside the tree trunk.
Dendroculus 5
It’s under a rock which needs to be hit by a dendro skill to be raised.
Dendroculus 6
Under the bridge.
Dendroculus 7
Dendroculus 8
Dendroculus 9
Just walk over the rocks to get this Dendroculus.
Dendroculus 10
It’s inside a cave. The entrance is south from Gandharva Ville.
Dendroculus 11
To dispel the field around it, you must shoot the floating targets with the dendrograna.
Dendroculus 12
Dendroculus 13
Just climb the tree so it will become visible.
Dendroculus 14
To access the area you must first interact with the flowers around it and then defeat the monsters.
Dendroculus 15
Dendroculus 16
It’s inside the mountain. The entrance on the right side of the giant robot.
Dendroculus 17
It has a forcefield protecting it which can be dispelled by using the harp you receive from the Aranyaka Quest.
Mawtiyima Forest and The Palace of Alcazarzaray Dendroculus locations in Genshin Impact
Below lies all of the Dendroculus locations we’ve located in Mawtiyima Forest and The Palace of Alcazarzaray in Genshin Impact:
Dendroculus 18
Dendroculus 19
Dendroculus 20
Dendroculus 21
Dendroculus 22
It’s pretty high in the air, so find a close place to jump from.
Dendroculus 23
It’s below the treetop, so use a Four-Leaf Sigil to reach it.
Dendroculus 24
Use the dendrograna to dispell the forcefield.
Dendroculus 25
Complete the Final Chapter Agnihotra Sutra World Quest to open the path.
Dendroculus 26
Dendroculus 27
Dendroculus 28
Climb the main building to the top.
Dendroculus 29
Next to a waterfall south from the Palace of Alcazarzaray.
Dendroculus 30
It’s inside a cave. The entrance is southwest from Mawtiyima Forest. You will find the Dendroculus at the top of a plant.
Devantaka Mountain West side Dendroculus locations in Genshin Impact
Here you’ll find all the Dendroculus locations we’ve uncovered on Devantaka Mountain in Genshin Impact:
Dendroculus 31
Inside a tree trunk.
Dendroculus 32
Dendroculus 33
In the giant robot’s index finger.
Dendroculus 34
On the giant robot’s face.
Dendroculus 35
Beneath the giant robot’s arm.
Dendroculus 36
Dendroculus 37
It’s beneath a rock, so use a dendro skill to lift it.
Dendroculus 38
At the top of the hill inside a shack.
Dendroculus 39
You need the song you learn in the Vimana Agam Quest.
Dendroculus 40
Port Ormos Dendroculus locations in Genshin Impact
Here are all the Dendroculus locations we’ve found in Port Ormos in Genshin Impact:
Dendroculus 41
Dendroculus 42
Dendroculus 43
Inside a tree trunk.
Dendroculus 44
Inside a tree trunk.
Dendroculus 45
Apam Woods Dendroculus locations in Genshin Impact
Below lies all of the Dendroculus locations we’ve found in Apam Woods in Genshin Impact:
Dendroculus 46
You must clear the Withering Zone to get it.
Dendroculus 47
At the top of a tree.
Dendroculus 48
You get access to it by progressing through the Varuna Gatha Quest.
Dendroculus 49
It’s beneath the tree roots.
Dendroculus 50
Dendroculus 51
Under a rock – you can access it by clearing the Withering Zone.
Dendroculus 52
Dendroculus 53
Dendroculus 54
Dendroculus 55
Dendroculus 56
It’s under a rock and to raise it you must clear the Withering Zone.
Dendroculus 57
It’s hidden under a pier of wood.
Dendroculus 58
It’s under a rock, so you must use a Dendro skill to raise it.
Dendroculus 59
Vimara Village and Pardis Dhyai Dendroculus locations in Genshin Impact
Here you’ll find all of the Dendroculus locations in Vimara Village and Pardis Dhyai in Genshin Impact:
Dendroculus 60
Dendroculus 61
Go south from Vimara Village.
Dendroculus 62
Near to Dendroculus 58.
Dendroculus 63
It’s under a rock. You need to clear the Withering Zone first to raise it.
Dendroculus 64
It’s at the top of a plant inside a cave – the entrance is north near the water.
Dendroculus 65
It’s inside a cave. To find the entrance, you need to swim from the large water area north of Pardis Dhyai.
Dendroculus 66
Dendroculus 67
Dendroculus 68
Dendroculus 69
Dendroculus 70
Dendroculus 71
It’s in a hidden entrance in the water level.
Dendroculus 72
Vanarana Dendroculus locations in Genshin Impact
Below you’ll find all of the Dendroculu locations in Vanarana in Genshin Impact:
Dendroculus 73
Dendroculus 74
Dendroculus 75
Dendroculus 76
To reach it, you must use a Dentro skill to activate the totems and drain the water in the area.
Dendroculus 77
It’s near to Dendroculus 72.
Dendroculus 78
Dendroculus 79
Dendroculus 80
Dendroculus 81
It’s inside a giant tree trunk.
Dendroculus 82
To find the house where the Dendroculus is, you must follow a route from the center of Vanarana.
Dendroculus 83
It’s under a rock, so you need to clear the Withering Zone first to raise it.
Dendroculus 84
Dendroculus 85
Sumeru City and Chatrakam Cave Dendroculus locations in Genshin Impact
Below you’ll find all of the Dendroculus locations in Sumeru City and Chatrakam Cave in Genshin Impact:
Dendroculus 86
It’s in the highest branch of the city’s tree.
Dendroculus 87
Dendroculus 88
You can only get it after progressing through the Agnihotra Sutra Quest.
Dendroculus 89
To find this one, you must activate the mushroom with dentro and use electro in the other mechanism. Use the Four-Leaf Sigils to reach it.
Dendroculus 90
You need to progress through the Agnihotra Sutra Quest to enter the cave.
Dendroculus 91
In the same cave as Dendroculus 86. Need to shoot some targets with dendrogranas to get it.
Dendroculus 92
Ruins of Dahri Dendroculus locations in Genshin Impact
Here you’ll find all of the Dendroculus locations for the Ruins of Dahri in Genshin Impact:
Dendroculus 93
It’s in a branch of an old tree near to a small arena.
Dendroculus 94
You need to progress through the Varuna Gatha Quest to find it.
Dendroculus 95
You need to progress through the Varuna Gatha Quest to find it.
Dendroculus 96
It’s hidden under a rock. To raise it, clear the Withering Zone first.
Dendroculus 97
To find it, you need to activate the portal by playing the Rhythm of the Gloomy Path song.
Dendroculus 98
Yasna Monument and Old Vanarana Dendroculus locations in Genshin Impact
Below lies all of the Dendroculus locations for Yasna Monument and Old Vanarana in Genshin Impact:
Dendroculus 99
You need to play the Rhythm of the Gloomy Path song to activate the portal.
Dendroculus 100
You need to progress through the quest Dream Nursery to open the path.
Dendroculus 101
It’s under a rock. Clear the Withering Zone to raise it.
Dendroculus 102
Dendroculus 103
Shoot the targets with the dendrograna to dispel the field around it.
Dendroculus 104
Dendroculus 105
Dendroculus 106
It’s under a rock. Use a Dendro skill to raise it.
Dendroculus 107
Dendroculus 108
You need to progress through the Aranyaka Quest to reach this region.
Dendroculus 109
Dendroculus 110
Aaru Village and Dar al-Shifa Dendroculus locations in Genshin Impact
Below lies all of the Dendroculus locations we’ve found so far in Aaru Village and Dar al-Shifa in Genshin Impact:
Dendroculus 111
Dendroculus 112
Dendroculus 113
Dendroculus 114
Near to a chest.
Dendroculus 115
Dendroculus 116
It’s inside a cave which you access by executing a plunging attack and hitting where the previous Ddendroculus was.
Dendroculus 117
Abdju Pit and West of Aaru Village Dendroculus locations in Genshin Impact
Here are all the Dendroculus locations we’ve found so far which lie west of Aaru Village and near Adju Pit in Genshin Impact:
Dendroculus 118
You’ll access this location during the Golden Slumber World Quest.
Dendroculus 119
You need to activate the elevator with the Scarlet Sand Slate which you gain during the Golden Slumber World Quest.
Dendroculus 120
You need to complete the Golden Slumber World Quest to activate with the Scarlet Sand Slate a new elevator in the same room where Ddendroculus 118 was.
Dendroculus 121
You need to complete the Golden Slumber World Quest to activate an elevator in the ruins you access in the Abdju Pit.
Dendroculus 122
You’ll find it during the Golden Slumber World Quest.
Dendroculus 123
To find it you need to access the ruins from the west and follow the path until you find a room with a time challenge. Complete it and use the elevator to find the room with the dendroculus.
Dendroculus 124
Dendroculus 125
Eyes of the Sands/Khaj-Nisut Dendroculus locations in Genshin Impact
Below are all of the Dendroculus locations we’ve found so far at the Eyes of the Sands – which, after you’ve completed a certain task, will become known as Khaj-Nisut – in Genshin Impact:
Dendroculus 126
Dendroculus 127
Dendroculus 128
You can access this building during the Golden Slumber World Quest.
Dendroculus 129
You can access this building during the Golden Slumber World Quest.
Dendroculus 130
Behind the rubble in one of the first rooms in the building.
Dendroculus 131
You access this building during the Golden Slumber World Quest. The room is turning left from where you get into the underground area.
Dendroculus 132
At the top of the building which only appears during the Golden Slumber World Quest.
Valley of Dahri Dendroculus locations in Genshin Impact
Here you’ll find all of the Dendroculus locations for the Valley of Dahri in Genshin Impact:
Dendroculus 133
Dendroculus 134
Dendroculus 135
Dendroculus 136
It’s in the lowest part of a cave where you can enter through a hole north from the Statue of the Seven.
Dendroculus 137
Dendroculus 138
It’s inside a temple which you can only access by using a specific password. To find it out, you must follow a marked NPC in the region.
Dendroculus 139
Dendroculus 140
It’s inside the Ruin Golem. You can get it during the Afratu’s Dilemma World Quest.
Dendroculus 141
Behind a giant gear inside the Ruin Golem. You can get it during the Afratu’s Dilemma World Quest.
Dendroculus 142
Dendroculus 143
Inside a cave that you can only access after completing the Afratu’s Dilemma World Quest.
Dendroculus 144
Inside a cave that you unlock after completing the Afratu’s Dilemma World Quest.
Hypostyle Desert Dendroculus locations in Genshin Impact
Below you’ll find all of the Dendroculus locations within the Hypostyle Desert region in Genshin Impact:
Dendroculus 145
Dendroculus 146
Dendroculus 147
Dendroculus 148
Dendroculus 149
At the top of the temple in the Dune of Elusion area.
Dendroculus 150
Dendroculus 151
You can find it during the Golden Slumber World Quest.
Dendroculus 152
You can find this one during the Golden Slumber World Quest.
Dendroculus 153
It’s in an underground area of the desert. You can access it during the Golden Slumber World Quest.
Dendroculus 154
This dendroculus is in an underground area of the desert. You can access it during the Golden Slumber World Quest.
Dendroculus 155
It’s also in the underground area of the desert. Keep up with the Golden Slumber World Quest to find it.
Dendroculus 156
You can find this room during the Golden Slumber World Quest.
Dendroculus 157
In one of the final areas you visit during the Golden Slumber World Quest.
Dendroculus 158
It’s inside at the end of a ruin which you can access it using an entrance in another ruin south from where the dendroculus is.
Dendroculus 159
Dendroculus 160
Dendroculus 161
Dendroculus 162
It’s inside the building. You must interact with the mechanism to find it.
Dendroculus 163
It’s in the middle of the air. To reach it, you must fly using a wind current located east from the dendroculus.
Dendroculus 164
Dendroculus 165
To enter the temple, you must go through a hole north from the teleport. Follow the path until you find a time challenge which you must complete to find the dendroculus.
Dendroculus 166
Dendroculus 167
Dendroculus 168
It’s in the underground part of the desert. You’ll find it during the Old Notes and New Friends World Quest.
Dendroculus 169
It’s inside the Dune of Magma which you can access during the Old Notes and New Friends World Quest.
Dendroculus 170
This one is in the underground area that you enter through the Dune of Magma during the Old Notes and New Friends World Quest.
Dendroculus 171
Keep following the area you found the previous dendroculus. This one is at the top of a giant mushroom.
Dendroculus 172
It’s in the middle of a withering zone in the underground area. It’s easy to get there by taking the southwest teleport in the Mausoleum of King Deshret.
Dendroculus 173
It’s easier to reach this one by using the southwest teleport in the Mausoleum of King Deshret.
Dendroculus 174
The entrance for the ruins is east from where the dendroculus is. To find it, you need to have completed part of the Old Notes and New Friends World Quest.
Dendroculus 175
It’s in the middle of the air, but to reach it you must have completed part of the Old Notes and New Friends World Quest.
Dendroculus 176
You can unlock it after completing the challenge next to where it is.
Dendroculus 177
Must have completed part of the Old Notes and New Friends World Quest to unlock the path to it.
Dendroculus 178
You will find it during the Old Notes and New Friends World Quest.
Dendroculus 179
You can find it by following the Old Notes and New Friends World Quest.
Dendroculus 180
It’s in the middle of the air in a room you find by doing the Old Notes and New Friends World Quest. To reach it, you need to activate the two Anemo totems in the room to start the wind current.
Desert of Hadramaveth Dendroculus locations in Genshin Impact
Below you’ll find all of the Desert of Hadramaveth Dendroculus locations in Genshin Impact:
Dendroculus 181
Dendroculus 182
It’s below a broken floor.
Dendroculus 183
Dendroculus 184
Dendroculus 185
At the entrance of a cave in the lower part in The Temples Forsaken area. Glide from the teleport.
Dendroculus 186
Further deep in the same cave as Dendroculus 185, above a group of enemies.
Dendroculus 187
It’s inside the temple, activate the mechanism outside to open the door.
Dendroculus 188
Follow ‘The Dirge of Bilqis’ World Quest to find it.
Dendroculus 189
Follow ‘The Dirge of Bilqis’ World Quest to find it.
Dendroculus 190
Follow ‘The Dirge of Bilqis’ World Quest to find it.
Dendroculus 191
Follow ‘The Dirge of Bilqis’ World Quest to find it.
Dendroculus 192
Dendroculus 193
Dendroculus 194
Dendroculus 195
Dendroculus 196
You access this area by following the World Quest The Dirge of Bilqis.
Dendroculus 197
Closer to the big robot you visit during the World Quest The Dirge of Bilqis.
Dendroculus 198
Dendroculus 199
You access this area by following the World Quest The Dirge of Bilqis.
Dendroculus 200
You can find it during the World Quest The Dirge of Bilqis Dune-Entombed Fecundity: Part II.
Dendroculus 201
Dendroculus 202
It’s found during the World Quest The Dirge of Bilqis Dune-Entombed Fecundity: Part III.
Dendroculus 203
It’s found during the World Quest The Dirge of Bilqis, but you need to use the flower mechanism to Activate Memory so the d Dendroculus becomes exposed.
Dendroculus 204
It’s found during the World Quest The Dirge of Bilqis The Eternal Dream, Ever Lush.
Dendroculus 205
It’s found during the World Quest The Dirge of Bilqis The Eternal Dream, Ever Lush.
Dendroculus 206
Inside a sandstorm. Must complete the World Quest The Dirge of Bilqis first to upgrade Jinni.
Dendroculus 207
Enter the underground area through a passage west of Dunes of Steel. Go south until you find a time challenge. Follow the path from it until a higher area where you’ll find rocks that can be destroyed. The Dendroculus is hiding behind it.
Dendroculus 208
First you must enter in the same underground tunnels you were in for Dendroculus 207. Walking south from the City of the Deceased domain, you must go south and drop in a hole opened once you interact with a rockpile. Go north now to a higher area which is in the same direction as the domain. The Dendroculus is hidden inside a hole on the wall.
Dendroculus 209
It’s in the entrance to the boss area.
Dendroculus 210
Dendroculus 211
It’s behind a well at the end of a Fatui camp.
Dendroculus 212
Dendroculus 213
Dendroculus 214
South of Debris of Panjhave.
Dendroculus 215
Dendroculus 216
Hidden behind a rock that must be destroyed by using a dendrogram.
Dendroculus 217
Close to a Shrine of Depth in the middle of the air.
Dendroculus 218
Close to a teleport north from The Sands of Three Canals.
Dendroculus 219
Dendroculus 220
Must activate the Anemo Totem to fly up to an invisible platform that takes you to the Dendroculus.
Dendroculus 221
In an area below the teleport.
Dendroculus 222
Dendroculus 223
Must first activate four dendro totems which are hidden in stones to activate a wind current for you to reach the Dendroculus. Break them using the dendrogram to find the totems.
Dendroculus 224
It’s hidden in a rock which must be destroyed by using dendrogram.
Dendroculus 225
It’s hidden in a rock which must be destroyed by using dendrogram.
Dendroculus 226
Dendroculus 227
Dendroculus 228
Dendroculus 229
You must use an invisible bridge to reach it in the middle of the air. Use the hilichurls as reference of where the bridge begins.
Dendroculus 230
It’s hidden in a rock which must be destroyed with a dendrogram. Must have access to Jinni to complete it.
Dendroculus 231
In the Giant’s eye.
Dendroculus 232
In the underground area you access through the World Quest The Dirge of Bilqis.
Dendroculus 233
Dendroculus 234
It’s beside a headless statue in the underground area.
Dendroculus 235
Accessible only during the World Quest Apocalypse Lost. Climb the statues, then the wall, to reach an invisible bridge where the Dendroculus is.
Gavireh Lajavar and Realm of Farakhert Dendroculus locations in Genshin Impact
Below you’ll find all of Dendroculus located in Gavireh Lajavar and Realm of Farakhkert regions in Genshin Impact:
Dendroculus 236
At the top of a hut west from the first statue you find.
Dendroculus 237
In the area above the domain.
Dendroculus 238
Dendroculus 239
Inside a cave unlocked during the ‘Khvarena of Good and Evil’ World Quest.
Dendroculus 240
At the top of a crystal in the Temri Mountains.
Dendroculus 241
It is easier to reach this dendroculus after the ‘Khvarena of Good and Evil’ world quest started.
Dendroculus 242
Dendroculus 243
You need start the ‘Khvarena of Good and Evil’ World Quest to have the ability to move the four-leaf sigil.
Dendroculus 244
Dendroculus 245
Dendroculus 246
Dendroculus 247
You will find it at the top of a crystal during the ‘Khvarena of Good and Evil’ World Quest. Use Sorush to complete a short challenge and unlock a new four-leaf sigil close to the Dendroculus.
Dendroculus 248
You will find it during the ‘Khvarena of Good and Evil’ World Quest.
Dendroculus 249
At the top of one of the giant roots in Vourukasha Oasis.
Dendroculus 250
Inside a stump near the oasis.
Dendroculus 251
Dendroculus 252
Dendroculus 253
You must use Sorush to open the green platform, destroy one of the orange stones closeby, and take the four-leaf sigil next to the Dendroculus.
Dendroculus 254
Use a dentro skill in a plant near the location which will unlock a four-leaf sigil right where the Dendroculus is.
Dendroculus 255
Dendroculus 256
It’s under the bridge.
Dendroculus 257
Dendroculus 258
At the top of a tent.
Dendroculus 259
At the top of the giant tree.
Dendroculus 260
Dendroculus 261
Inside a structure of metal.
Dendroculus 262
Dendroculus 263
You can use Sorush to place a four-leaf sigil next to this one.
Dendroculus 264
This one is inside a cave you can enter by using three different entrances. To find it, you must first destroy some rocks close to a Hilichurl digging the ground to find a Farrwick that will open the path blocked by gray crystals.
Dendroculus 265
This one is underground. During the ‘Khvarena of Good and Evil’ World Quest, after you solve the puzzle, you can use an elevator in the same room to go to the upper floor where you will find the Dendroculus right in front of you.
Dendroculus 266
It’s found during the ‘Khvarena of Good and Evil’ World Quest in the ‘As the Khvarena’s Light Shows’ segment.
Dendroculus 267
Since it’s underground, you must follow the ‘Khvarena of Good and Evil’ World Quest to get it.
Dendroculus 268
The area where you can find this one is underground and accessible through the ‘Khvarena of Good and Evil’ World Quest. Once you open the first door in the puzzle closer to the Tunigi Hollow teleport, go to the right and use the wind torrent to reach the pipes where this Dendroculus is.
Dendroculus 269
It’s underground, above a furnace you will find during the ‘Khvarena of Good and Evil’ World Quest.
Dendroculus 270
Under the elevator you unlock after a puzzle during the ‘Khvarena of Good and Evil’ World Quest.
Dendroculus 271
Floating above the water. Use Sorush to take the four-leaf sigil close to it.
And these are all the Dendroculus you can find in Genshin Impact. You can also check out our guides on the first Archon Quest for the Sumeru region – Through Mists of Smoke and Forests – and how to unlock Sumeru Reputation.
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